Between Wellfleet and Provincetown the Cape narrows to a strip, in places barely more than a mile from the Atlantic to the bay, and that strip is Truro. It has 2,454 year-round residents and no downtown. There is a post office, a small cluster at Truro Center by the Pamet River, another at North Truro, and otherwise hills, moor, pine, dune and a great deal of federal land. More than half the town's area sits inside the Cape Cod National Seashore.
This is either the best or the worst place on the Cape to stay, depending entirely on what you came for. If you want to walk out of a rental and buy a coffee, it is the worst. If you want the emptiest ocean beaches in Massachusetts and a sky with nothing in it, there is nowhere better.
The landscape here is unlike the rest of the peninsula. Truro's interior rolls. The glacial outwash has been carved into steep hollows and hills by wind and by the Pamet, a tidal river cutting almost the full width of the Cape from the bay side. Walk the Pamet valley trails and you are in a wet, green, enclosed landscape that could not look less like a dune. Ten minutes later you are on top of a bluff with the Atlantic two hundred feet below.
The Wampanoag called this ground Pamet, or Payomet, and it is where the Mayflower party first came ashore in November 1620, before they went on to Plymouth. At Corn Hill they dug up a cache of stored seed corn, which is why the hill is called that and why the story is more complicated than the version most Americans were taught.
Two things dominate the skyline. Highland Light, the oldest and tallest lighthouse on the Cape, and the radar tower on the old North Truro Air Force Station, now part of the Seashore. Beyond that, there is very little built. That is the entire proposition.